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Feral pigeon

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A large flock of feral pigeons

Feral pigeons (Columba livia domestica or Columba livia forma urbana), also called city doves, city pigeons, or street pigeons,[1][2] are descendants of domestic pigeons (Columba livia domestica) that have returned to the wild.[3] The domestic pigeon was originally bred from the wild rock dove, which naturally inhabits sea-cliffs and mountains.[4] Rock, domestic, and feral pigeons are all the same species and will readily interbreed. Feral pigeons find the ledges of buildings to be a substitute for sea cliffs, have become adapted to urban life, and are abundant in towns and cities throughout much of the world.[5]

Owing to their capacity to create large amounts of excrement and be an occasional disease vector to humans combined with crop and property damage, pigeons are largely considered a nuisance and an invasive species,[6][7] often disparagingly referred to as "rats with wings".[8][9][10] Actions are taken in many municipalities to lower their numbers or completely eradicate them.[1][2][11][12][13][14]

Feral pigeons are considered invasive in many parts of the world,though they have a positive impact on wild bird populations, serving as an important prey species for birds of prey.


Physical characteristics

[edit]
Main article: Rock dove § Description

Feral pigeons are essentially the same size and shape as the original wild rock dove, but often display far greater variation in colour and pattern than their wild ancestors. The blue-barred pattern which the original wild rock dove displays is generally less common in more urban areas. Urban pigeons tend to have darker plumage than those in more rural areas.[15]

  • White
    White
  • Brown
    Brown
  • Black
    Black
  • Pied
    Pied

Genetics

[edit]

The avoidance of mating between related individuals is ordinarily regarded as adaptive since it decreases the likelihood of inbreeding depression in progeny that can be caused by the expression of deleterious recessive alleles.[16] However in feral pigeons it was found that despite detectable inbreeding depression, pairwise relatedness between mates was significantly greater than it was between nonmates.[16] This suggests that mating with close kin provides inclusive fitness benefits that outweigh the costs of inbreeding depression.[16]

Protection status

[edit]

In the United Kingdom, pigeons are covered under the "General Licences" and can be humanely culled by the land owner or their agent for a variety of reasons including spread of human disease. It is illegal to kill/destroy nests for any reason other than those listed under the general licences.[discuss]

In the U.S., the Migratory Bird Treaty Act of 1918, which protects native birds, does not apply to feral pigeons, common starlings or house sparrows, because they are introduced species.[17] It is usually legal to kill feral pigeons in the United States; methods such as poisons may be regulated, however.[18] Pigeons are labeled an invasive species in North America by the USDA.[14]

In India, pigeons are protected under Section 428 and Section 429 of the Indian Penal Code. Wild pigeons are further protected under the Wildlife Protection Act, 1972.[19]

Impact

[edit]
A flock of feral pigeons at rest in front of the statue of James II, London

Feral pigeons often only have small populations within cities relative to the number of humans. For example, the breeding population of feral pigeons in Sheffield, England in summer 2005 was estimated at 12,130 individuals (95% confidence interval 7757–18,970), in a city with a human population of about 500,000.[20] Despite this, feral pigeons usually reach their highest densities in the central portions of cities, so they are frequently encountered by people, which may lead to conflict.

Potential health risk to humans

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Feral pigeons are widely considered pests, and can be reservoirs and vectors of some human and livestock diseases, such as salmonellosis and tuberculosis.[21][22][23] However, it is rare that a pigeon will transmit a disease to humans due to their immune system.[23] Although feral pigeons pose sporadic health risks to humans, the risk is low, even for humans involved in occupations that bring them into close contact with nesting sites. Analysis revealed that feral pigeons harbored a total of 60 different human pathogenic organisms. Five pathogens were viruses, nine were bacteria, 45 were fungi, and one was a protozoan. However, only five pathogens were routinely transmitted to humans. There were single case incidences for transmission of Salmonella enterica. [24]

Pigeon feces has been implicated as a "contributing factor" a cryptococcus infection leading to the death of a child.[25]

Feces

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Healthy, well-fed pigeons excrete feces that are composed largely of a brown to greenish-brown mass with a white cap; the white cap is composed of uric acid crystals, with some of the uric acid dissolved within urea that comprises the liquid urine, which is transparent and often absorbed by the mass of solid waste of the feces, which maintains its shape when the bird is in good health. The color of the resultant fecal matter is helpful in diagnosing the condition of the bird; droppings which appear dull yellow is indicative of starvation especially if watery, green may be a sign of liver disease due to an abundance of excreted biliverdin, bright green being an abundance of passed bile implying bowel problems or not having eaten recently, and red-brown showing kidney disease.[26]

It is often claimed that feral pigeon feces cause substantial economic losses from property damage, often said to be from their corrosive droppings damaging stone edifices and metal structures.[27][28][29][30]

Control methods

[edit]
See also: Animal population control
Many places where feral pigeons could land are covered with spikes.

Deterrence methods include bird spikes and the recently introduced optical gel deterrent.[31]

Predators

[edit]

Peregrine falcons, which are also originally cliff dwellers, have too adapted to the skyscrapers of large cities and often feed exclusively on rock doves.[32] Some cities actively encourage this through falcon breeding programs. Projects include the Unibase Falcon Project and the Victorian Peregrine Project.

Other predators of the pigeon have been recorded, including Eurasian sparrowhawks, crows and gulls.[33][34][35] In London, the population of great white pelicans at St. James's Park has also been recorded killing and consuming pigeons even when alternative food sources are available.[36][37] In cities in Western Europe, European herring gulls may occasionally hunt and consume feral pigeons in addition to other birds and small mammals.[38] Goshawks have been known to specialize on feral pigeons even outside of urban settings.[39][40][41]

Larger birds of prey occasionally take advantage of this population as well. In New York City, the abundance of feral pigeons (and other small animals) has created such a conducive environment for predators that the red-tailed hawk has begun to return in very small numbers, including the notable Pale Male.[42]

Despite their importance in the diet of urban predatory birds, feral pigeons are known to transmit several diseases to their consumers, as they may be reservoirs of several avian diseases, often subclinically. Raptors may contract West Nile Virus through consumption of infected bird species, such as crows, house sparrows, and pigeons.[43] Trichomoniasis, frounce, or canker is a significant disease in raptors both captive and wild-living. Symptoms include caseous plaques in and around the oropharynx, and lesions may be found in the lungs, air sacs, sinuses, ear canal and kidneys.[43] The lesions may progress into severe necroses, at least of the upper digestive tract.[44] Trich has been found to be resistant to several drug treatments, and it is thought that "preventive treatments" that use insufficient dosages may be one of the causes.[45] These diseases may prove to be fatal to the raptors; Columbid Herpesvirus-1 has a near 100% mortality rate in raptors.[46]

Poison

[edit]

Due to their non-selective nature, most avian poisons have been banned. In the United States market, only 4-aminopyridine (Avitrol) and DRC-1339 remain registered by EPA. DRC-1339 is limited to USDA use only, while 4-AP is a restricted-use pesticide, for use only by licensed applicators.

The use of poisons has been proven to be fairly ineffective, however, as pigeons can breed very quickly, and their numbers are determined by how much food is available; that is, they breed more often when more food is provided to them. When pigeons are poisoned, surviving birds do not leave the area. On the contrary, they are left with more food per bird than before. This attracts pigeons from outside areas as well as encouraging more breeding, and populations are re-established quickly.[47] An additional problem with poisoning is that it also kills pigeon predators. Due to this, in cities with peregrine falcon programs it is typically illegal to poison pigeons.[32]

Reducing food supply

[edit]
A man feeding feral pigeons at Esplanadi in Helsinki, Finland in 1921
One of the difficulties of controlling feral pigeon populations is the common practice of feeding them, as here in New York City.
Video showing feral pigeons eating seeds

A more effective tactic to reduce the number of feral pigeons is deprivation.[48] Cities around the world have discovered that not feeding their local birds results in a steady population decrease in only a few years. As scavengers, pigeons will still pick at garbage bags containing discarded food or at leftovers carelessly dropped on the ground, but securely disposing of foodstuffs will greatly reduce scavenger populations. Feeding of pigeons is banned in parts of Venice, Italy.[49]

Long-term reduction of feral pigeon populations can be achieved by restricting food supply, which in turn involves legislation and litter (garbage) control. Some cities have deliberately established favorable nesting places for pigeons—nesting places that can easily be reached by city workers who regularly remove eggs, thereby limiting their reproductive success.[2] In addition, pigeon populations may be reduced by bird control systems that successfully reduce nesting sites.

Avian contraceptives

[edit]
See also: Wildlife contraceptive

In 1998, in response to conservation groups and the public interest, the National Wildlife Research Center (NWRC), a USDA/APHIS laboratory in Fort Collins, Colorado, started work on nicarbazin, a promising compound for avian contraception. Originally developed for use in resident Canada geese, nicarbazin was introduced for use as a contraceptive for feral pigeons in 2007.

The active ingredient, nicarbazin, interferes with the viability of eggs by binding the ZP-3 sperm receptor site in the egg.[50] This unique contraceptive action is non-hormonal and fully reversible.[51]

Registered by the EPA as a pesticide (EPA Reg. No. 80224-1), "OvoControl P", brand of nicarbazin, is increasingly used in urban areas and industrial sites to control pigeon populations. Declared safe and humane, the new technology is environmentally benign[52] and does not represent a secondary toxicity hazard to raptors or scavengers.[53]

Avian contraception has the support of a range of animal welfare groups including the Humane Society of the United States, the American Society for the Prevention of Cruelty to Animals and People for the Ethical Treatment of Animals. Avian contraceptives are also perceived by some civilians as an acceptable method for population control, over other methods such as prohibition to feeding or extermination.[54]

Dummy egg nesting

[edit]
Large pigeon trap/coop/loft at Batman Park, Melbourne. Designed specifically to encourage nesting and allow removal of fertilised eggs to prevent population growth, it was a landmark in its own right before its removal, for lack of success, around 2015.

When eggs are removed in artificial pigeon houses, the interval between reproductive attempts is strongly reduced, which reduces the efficiency of the method.[55] Dummy egg nesting programs have therefore been tested in some cities with mixed results. There, the eggs are removed and replaced with dummy eggs. The real eggs are then destroyed. One such structure, in Batman Park in Melbourne, Australia, was unsuccessful in attracting pigeons and has since been removed.[56][57] The loft used in Melbourne was on stilts, with a cage door allowing access from beneath for accessing the structure at night when the pigeons are asleep.

Monitoring pigeon population

[edit]

Estimating the population size of pigeons is necessary for monitoring and control programs of pigeons in parks and other urban areas. The methods used for estimating populations sizes are:

  • Stratified grids: This method consists in dividing the area where pigeons occur in 500x500m squares. 34% of the squares are selected randomly and pigeons are counted in a 5 meters radius for 5 minutes.[58][59]
  • Point-counts: standing in the center of a park, the observer makes a 360 degree turn while counting individuals with a manual mechanical counter in a radius of approximately 50m, limited by the streets and buildings that surround the park.[60]
  • Panoramas: taking 360 panoramic photographs, while standing at the center of the park, and using software to place a number above the counted pigeon in the panoramic photograph. This method has been proven the most effective of all.[61]

City squares famous for pigeons

[edit]

Many city squares have large pigeon populations, such as Washington Square Park in New York City, George Square in Glasgow, the Piazza San Marco in Venice, Dam Square in Amsterdam, The Gateway of India and Kabutarkhana in Mumbai and (prior to 2000) Trafalgar Square in London.[11]

Pigeon by artist Paul Sloan at Rundle Mall, Adelaide

A 2 metres (6 ft 7 in) tall statue of a pigeon by artist Paul Sloan was installed at the Rundle Mall, Adelaide, South Australia, adding to their collection of art installations, including statues of pigs. Sloan intended to "elevate the humble pigeon" with his work titled Pigeon. The mirrored stainless steel statue cost AU$174,000. While the installation has been talked up by City of Adelaide Lord Mayor Sandy Verschoor, some locals have responded negatively.[62][63]

See also

[edit]
  • Bird feeding – typically thought of as an activity of bird enthusiasts, studies have revealed it may have both positive and negative impact
  • Doves as symbols – appearing, usually white in color, in many settings as symbols of love, peace or as messengers, in the symbolism of various religions and of both military and pacifist groups
  • Squab – a young pigeon, typically under four weeks old, or its meat
  • Feral chicken
  • Feral parrot

References

[edit]
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Books

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